Drake, A.G.; et al.; Losos, J.B.. Proceedings of the National Academy of Sciences, 2025
PNAS 122(18):e2413780122, 28 Apr 2025.
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BOAS involves narrowed nostrils, an overlong soft palate and an undersized trachea, and pugs are among the breeds most affected.
The American Kennel Club recognized the pug in 1885.
A popular belief derives the name 'pug' from the Latin pugnus, meaning fist.
Pugs are prone to hemivertebrae.
A UK VetCompass study of veterinary records found pugs have a life expectancy of about 7.7 years, versus about 11.2 for dogs in general.
An April 2025 PNAS study led by Cornell's Abby Grace Drake found that pugs and Persian cats have converged on skull shapes more similar to each other than to their wild ancestors.
Konrad Lorenz proposed the baby schema: infant-like features that trigger caregiving responses.
About 40% of pugs carry the high-risk marker for pug dog encephalitis and 11% carry two copies. The marker test was developed at UC Davis in 2010.
Pug dog encephalitis (necrotizing meningoencephalitis) is a fatal inflammatory brain disease.
Drake, A.G.; et al.; Losos, J.B.. Proceedings of the National Academy of Sciences, 2025
PNAS 122(18):e2413780122, 28 Apr 2025.
O'Neill, Dan G.; Darwent, Elisabeth C.; Church, David B.; Brodbelt, Dave C.. Canine Genetics and Epidemiology, 2016
Teng, Kendy Tzu-yun; Brodbelt, Dave C.; Pegram, Camilla; Church, David B.; O'Neill, Dan G.. Scientific Reports, 2022
Glocker, Melanie L.; Langleben, Daniel D.; Ruparel, Kosha; et al.. PNAS, 2009
Archived copy (Wayback Machine).
American Kennel Club
Purina Pro Club
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